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In the next 24 hours, this layer allows the process of bacterial adhesion to occur, with both diatoms and bacteria (e.g.

vibrio alginolyticus, pseudomonas putrefaciens) attaching, initiating the formation of a biofilm.

Fouling detection relies on the biomass' property of fluorescence.

All microorganisms contain natural intracellular fluorphores, which radiate in the UV range when excited.

By the end of the first week, the rich nutrients and ease of attachment into the biofilm allow secondary colonizers of spores of macroalgae (e.g. Within 2 to 3 weeks, the tertiary colonizers- the macrofoulers- have attached.

Biofouling is divided into microfouling — biofilm formation and bacterial adhesion — and macrofouling — attachment of larger organisms.Industrially important impacts are on the maintenance of mariculture, membrane systems (e.g., membrane bioreactors and reverse osmosis spiral wound membranes) and cooling water cycles of large industrial equipment and power stations.Biofouling can occur in oil pipelines carrying oils with entrained water, especially those carrying used oils, cutting oils, oils rendered water-soluble through emulsification, and hydraulic oils.Examples of non-calcareous (soft) fouling organisms are seaweed, hydroids, algae and biofilm "slime".Marine fouling is typically described as following four stages of ecosystem development.At UV-range wavelengths, such fluorescence arises from three aromatic amino acids - tyrosine, phenylalanine, and tryptophan.